[1] Univ Carthage, Fac Sci Bizerte, Zarzouna, Tunisia
[2] Univ Bordeaux, UMR CNRS 5805 EPOC, Allee Geoffroy St Hilaire CS 50023, FR-33615 Pessac, France
[3] Ctr Biotechnol Bed Cedria, Lab Extremophiles Plants, Hammam Lif, Tunisia
[4] Lanzhou Univ, Sch Life Sci, State Key Lab Grassland & Agro Ecosyst, Lanzhou, Gansu, Peoples R China