A new species of ground beetle and a revised list of the East Asian endemic subgenus Nialoe(s. lat.) (Coleoptera: Carabidae: Pterostichus)

The taxonomy of the East Asian endemic subgenusNialoe(s. lat.) Tanaka, 1958 (Coleoptera: Carabidae:Pterostichus) is still insufficient at both the specific and subgeneric levels, and the discovery of new species and revision of the subgeneric classification are ongoing. This study escribes Pterostichus(Nialoe) hekosanensissp. nov. from central Honshu, Japan. The new species is similar toP. daihizanusIshida, 1968;P. ohbayashiiIshida, 1968; andP. ryoheiiMorita, 2012, but can be readily distinguished from them by distinct characters. In addition, the subgenusNialoein the last version of the Palaearctic Catalogue, which is widely accepted by experts, is reviewed from a phylogenetic perspective and a revised list reflecting the phylogenetic relationships is presented.