[1] Univ Antwerp, Ecosyst Management Res Grp, B-2610 Antwerp, Belgium
[2] Royal Netherlands Inst Sea Res, Spatial Ecol Res Grp, NL-4400 AC Yerseke, Netherlands
[3] Inst Marine Resources & Ecosyst Studies IMARES, NL-4400 AB Yerseke, Netherlands
[4] Delft Univ Technol, NL-3311 JG Dordrecht, Netherlands
[5] EcoShape, NL-3311 JG Dordrecht, Netherlands