[1] Osaka Univ, Grad Sch Frontier Biosci, Cognit Neurosci Lab, Toyonaka, Osaka 5608531, Japan
[2] Osaka Univ, Ctr Informat & Neural Networks CiNet, Suita, Osaka 5650871, Japan
[3] Natl Inst Informat & Commun Technol, Suita, Osaka 5650871, Japan
[4] Japan Sci & Technol Agcy, CREST, Kawaguchi, Saitama 3320012, Japan
[5] Univ Tokyo, Grad Sch Med, Dept Neurophysiol, Bunkyo Ku, Tokyo 1130033, Japan