[1] Shandong Agr Univ, State Key Lab Crop Biol, Shandong Key Lab Crop Biol, Coll Life Sci, Tai An 271018, Shandong, Peoples R China
[2] Univ Calif San Diego, Sect Cell & Dev Biol, La Jolla, CA 92093 USA
[3] Chinese Acad Sci, State Key Lab Plant Genom, Inst Genet & Dev Biol, Beijing 100101, Peoples R China
[4] Chinese Acad Sci, Natl Plant Gene Res Ctr, Inst Genet & Dev Biol, Beijing 100101, Peoples R China