[1] Dire Dawa Univ, Dept Biol, POB 13-62, Dire Dawa, Ethiopia
[2] Haramaya Univ, Coll Agr & Environm Sci, Sch Plant Sci, Genet & Plant Breeding Program, Dire Dawa, Ethiopia
[3] Haramaya Univ, Coll Agr & Environm Sci, Sch Plant Sci, Plant Protect Program, Dire Dawa, Ethiopia
[4] United States Agr Anim & Plant Hlth Inspect Serv, Plant Protect & Quarantine, Natl Identificat Serv, Riverdale, MD USA
[5] Ethiopian Inst Agr Res, Holetta Agr Res Ctr, Addis Ababa, Ethiopia