[1] Japan Sci & Technol Agcy, CREST, Kawaguchi, Saitama 3320012, Japan
[2] Hokkaido Univ, Inst Low Temp Sci, Kita Ku, Sapporo, Hokkaido 0600819, Japan
[3] Vrije Univ Amsterdam, Fac Earth & Life Sci, Dept Hydrol & Geoenvironm Sci, NL-1081 HV Amsterdam, Netherlands
[4] Ctr Ecol & Hydrol, Wallingford OX10 8BB, Oxon, England