[1] Seoul Natl Univ, Coll Vet Med, Microbiol, Seoul 151742, South Korea
[2] Seoul Natl Univ, BK21 Program Vet Sci, Seoul 151742, South Korea
[3] Seoul Natl Univ, Coll Vet Med, KRF Zoonot Dis Prior Res Inst, Seoul 151742, South Korea
[4] Konkuk Univ, Food Sci & Biotechnol Anim Resource, Seoul 143701, South Korea