[1] Shenyang Agr Univ, Coll Agron, Shenyang 110866, Peoples R China
[2] Chinese Acad Sci, Inst Geog Sci & Nat Resources Res, Key Lab Ecosyst Network Observat & Modeling, Beijing 100101, Peoples R China
[3] Chinese Acad Sci, Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[4] Univ Chinese Acad Sci, Coll Resources & Environm, Beijing 100190, Peoples R China
[5] Chinese Acad Sci, Northeast Inst Geog & Agroecol, Key Lab Wetland Ecol & Environm, Changchun 130102, Peoples R China