Scaphitid ammonites from the lower Maastrichtian of Nahoryany (western Ukraine) and discussion on the origin of Hoploscaphites constrictus

The lower Maastrichtian (Upper Cretaceous) section at Nahoryany, south of Lviv (western Ukraine), has been known as a rich source of fossils since 1843. However, the section is not available since a long time ago and fossils collected from it can only be studied in museum collections. In the present paper, ammonites of the scaphitid genus Hoploscaphites are studied based on Nahoryany material, comprising Hoploscaphites pumilus, H. constrictus and H. sp. Specimens from Nahoryany in museum collections lack data on their provenance level within the section so it cannot be determined whether or not they occurred together or formed a succession; data on inoceramid bivalves indicate the presence of two successive zones at Nahoryany. Some specimens from Nahoryany here referred to as H. pumilus are similar to the controversial H. constrictus anterior from the lower Maastrichtian of Poland. This morphology suggests an ancestor-descendant relationship between H. pumilus and H. constrictus, a common species once considered as an informal index for the Boreal Maastrichtian in Europe. The inferred evolutionary transition from H. pumilus to H. constrictus was towards a less robust, recoiled shell, which would have enhanced the horizontal swimming ability and manoeuvrability of these cephalopods. Recoiling trends occurred in several lineages of Late Cretaceous scaphitids.