[1] South China Agr Univ, Coll Agr, State Key Lab Conservat & Utilizat Subtrop Agro Bi, Guangzhou 510642, Peoples R China
[2] Yulin Normal Univ, Guangxi Key Lab Agr Resources Chem & Biotechnol, Guangxi 537000, Peoples R China
[3] Minist Agr & Rural Affairs, Sci Observing & Expt Stn Crop Cultivat South China, Guangzhou 510642, Peoples R China
[4] Guangzhou Key Lab Sci & Technol Fragrant Rice, Guangzhou 510642, Peoples R China