[1] Jeonbuk Natl Univ, Dept Agr Biol, Jeonju 54896, South Korea
[2] Chonnam Natl Univ, Inst Environmentally Friendly Agr, Dept Appl Biol, Gwangju 61185, South Korea
[3] Natl Inst Hort & Herbal Sci, Rural Dev Adm, Hort & Herbal Crop Environm Div, Wonju 55365, South Korea
[4] Jeonbuk Natl Univ, Dept Plant Protect & Quarantine, Jeonju 54896, South Korea
[5] Jeonbuk Natl Univ, Dept Agr Convergence Technol, Jeonju 54896, South Korea