[1] Harbin Normal Univ, Coll Life Sci & Technol, Harbin 150025, Peoples R China
[2] Univ Colorado, Museum Nat Hist, Boulder, CO 80309 USA
[3] Univ Colorado, Dept Ecol & Evolutionary Biol, Boulder, CO 80309 USA
[4] RAS, K A Timiryazev Inst Plant Physiol, IPP, Moscow 127276, Russia
[5] Shanghai Normal Univ, Coll Life Sci, Shanghai 200234, Peoples R China