A new putative moss burl (Insecta: Hemiptera) from the lower Permian of the Saar-Nape Basin, SW Germany, and the age of Coleorrhyncha

Permoridium fresenaci gen. et sp. nov., the oldest putative representative of the bug suborder Coleorrhyncha, is described from the lower Permian of Germany, in the new family Permoridiidae. The main autapomorphies of the new taxon are the separation of M+CuA from R at extreme wing base and the veins CuP and PCu strongly appressed. Its position in the Hemiptera is discussed. Its attribution to the modern Coleorrhyncha is supported by the putative synapomorphies 'a very long and well-defined ScP, ending into RA', 'ScP appressed R +M+CuA at extreme wing base' and 'a strong ScA, distant from costal vein and with a short anterior branch at its extreme base, ending into C, plus posterior branches'. The relationships of the extinct families Hoploridiidae, Karabasiidae and Progonocimicidae with the extant Coleorrhyncha are discussed.