[1] Univ Politecn Madrid UPM, Ctr Biotecnol & Genomica Plantas, Inst Nacl Invest & Tecnol Agr & Alimentac INIA CSI, Campus Montegancedo, Madrid 28223, Spain
[2] Univ Barcelona, Fac Biol, Dept Evolutionary Biol Ecol & Environm Sci, Barcelona 08028, Spain
[3] Univ Politecn Madrid UPM, Escuela Tecn Super Ingn Agron Alimentaria & Biosis, Dept Biotecnol Biol Vegetal, Madrid 28040, Spain