[1] Northeast Forestry Univ, Coll Wildlife & Protected Area, Harbin, Peoples R China
[2] First Hosp Jilin Univ, Key Lab Organ Regenerat & Transplantat, Key Lab Zoonot Dis, Dept Infect Dis,Minist Educ, Changchun, Peoples R China
[3] First Hosp Jilin Univ, Ctr Infect Dis & Pathogen Biol, Key Lab Organ Regenerat & Transplantat, Minist Educ Key,Lab Zoonot Dis, Changchun, Peoples R China
[4] Kansas State Univ, Coll Vet Med, Dept Diagnost Med Pathobiol, Manhattan, KS 66506 USA
[5] Chinese Acad Agr Sci, Harbin Vet Res Inst, State Key Lab Vet Biotechnol, Harbin, Peoples R China