[1] Univ Tecnol Bolivar, Parque Ind & Tecnol Carlos Velez Pombo Km 1, Cartagena 130010, Colombia
[2] Univ Cauca, Calle 5 4-70, Popayan 190001, Colombia
[3] Ctr Invest Oceanog Hidrograf Pacifico DIMAR, Capitania Puerto Tumaco, San Andres De Tumaco, Colombia
[4] Fdn Edmund Mach, Res & Innovat Ctr, Via E Mach 1, I-38098 San Michele All Adige, Italy