[1] Univ Oxford, John Radcliffe Hosp, Dept Clin Neurosci, Nuffield Lab Ophthalmol, West Wing, Oxford OX3 9DU, England
[2] Oxford Univ Hosp NHS Fdn Trust, John Radcliffe Hosp, Oxford Eye Hosp, Headley Way, Oxford OX3 9DU, England
[3] UCL Inst Ophthalmol, 11-43 Bath St, London EC1V 9EL, England
[4] Moorfields Eye Hosp, NIHR Moorfields Biomed Res Ctr, London, England