[1] Humboldt Univ, Dept Geog, Lab Landscape Ecol, Rudower Chaussee 16, D-12489 Berlin, Germany
[2] UFZ Helmholtz Ctr Environm Res, Dept Computat Landscape Ecol, Permoserstr 15, D-04318 Leipzig, Germany
[3] Univ Shef fi eld, Dept Urban Studies & Planning, Western Bank, Sheffield S10 2TN, England
[4] Ruhr Univ Bochum, Inst Geog, Universitatsstr 150, D-44801 Bochum, Germany