[1] Australian Museum, Australian Museum Res Inst, Sydney, NSW 2010, Australia
[2] Macquarie Univ, Dept Biol Sci, N Ryde, NSW 2109, Australia
[3] Univ Bordeaux, CNRS, Observ Aquitain Sci Univers, UMS 2567,POREA, Pessac, France
[4] Univ Sydney, Sch Life & Environm Sci, Camperdown, NSW 2006, Australia
[5] Univ Calif San Diego, Scripps Inst Oceanog, La Jolla, CA 92093 USA
[6] Univ Bordeaux, CNRS, Bordeaux INP, EPOC,UMR 5805, F-33120 Arcachon, France