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Yangzhou Univ, Jiangsu Key Lab Crop Genet & Physiol, Jiangsu Coinnovat Ctr Modern Prod Technol Grain C, Jiangsu Key Lab Crop Cultivat & Physiol,Agr Coll, 88 Daxue South Rd, Yangzhou 225009, Jiangsu, Peoples R China Yangzhou Univ, Jiangsu Key Lab Crop Genet & Physiol, Jiangsu Coinnovat Ctr Modern Prod Technol Grain C, Jiangsu Key Lab Crop Cultivat & Physiol,Agr Coll, 88 Daxue South Rd, Yangzhou 225009, Jiangsu, Peoples R China