[1] Iranian Res Org Sci & Technol IROST, Museum Iranian Lichens, Tehran, Iran
[2] Iranian Res Org Sci & Technol IROST, Dept Biotechnol, Tehran, Iran
[3] Brigham Young Univ, Dept Biol, Provo, UT 84602 USA
[4] Brigham Young Univ, M L Bean Life Sci Museum, Provo, UT 84602 USA
[5] Karl Franzens Univ Graz, Inst Biol, Div Plant Sci, NAWI Graz, A-8010 Graz, Austria