[1] Lanzhou Univ, Coll Pastoral Agr Sci & Technol, State Key Lab Herbage Improvement & Grassland Agro, Key Lab Grassland Livestock Ind Innovat,Minist Agr, Lanzhou 730000, Peoples R China
[2] Yunnan Univ, Inst Int Rivers & Ecosecur, Yunnan Key Lab Soil Eros Prevent & Green Dev, Kunming 650500, Peoples R China
[3] Lanzhou Univ Finance & Econ, Sch Agr & Forestry Econ & Management, Lanzhou 730020, Peoples R China
[4] Chinese Acad Sci, Northwest Inst Plateau Biol, Key Lab Cold Reg Restorat Ecol, Xining 810008, Qinghai, Peoples R China
[5] Qinghai Normal Univ, Dept Life Sci, Xining 810008, Peoples R China
[6] Yunnan Univ, Sch Ecol & Environm Sci, Kunming 650091, Peoples R China