[1] Univ Oxford, Peter Medawar Bldg Pathogen Res, Oxford, England
[2] Katholieke Univ Leuven, Univ Hosp Leuven, Dept Gastroenterol & Hepatol, Leuven, Belgium
[3] Univ Oxford, Translat GastroIntestinal Unit, Nuffield Dept Med, Oxford, England
[4] Icahn Sch Med Mt Sinai, Henry Janowitz Div Gastroenterol, New York, NY USA
[5] Univ Warwick, Warwick Med Sch, Div Biomed Sci, Coventry, England
[6] Univ Pennsylvania, Perelman Sch Med, Div Gastroenterol, Dept Med, Philadelphia, PA USA