[1] Osaka Univ, Grad Sch Engn Sci, 1-3 Machikaneyama Cho, Toyonaka, Osaka 5608531, Japan
[2] Natl Inst Adv Ind Sci & Technol, 1-1-1 Umezono, Tsukuba, Ibaraki 3058563, Japan
[3] Shiga Univ Med Sci, Dept Fundamental Biosci, Otsu, Shiga 5202192, Japan
[4] Osaka Univ, SANKEN Inst Sci & Ind Res, 8-1 Mihogaoka, Osaka, Ibaraki 5670047, Japan