A New Species of the Genus Nanorana Günther, 1896 (Anura: Dicroglossidae) from Hengduan Mountains of China

An integrative study based on morphological data and mitochondrial molecular data of all known Nanorana (sensu stricto) species have revealed a new species from the Hengduan Mountains, N. huangi sp. nov. The new species can be distinguished from its congers by its: (1) medium body size, SVL 31.3-40.6 mm in adult males (n = 12), 39.2-48.1 mm in adult females (n = 8); (2) absence of subarticular tubercles on fingers, with supernumerary tubercles below the base of the fingers small but distinct; (3) presence of dark dorsal patches without pale margins, generally creamy yellow ventral body basically, bright yellow ventral limbs; (4) small but prominent tympanum; (5) tibio-tarsal joint that reaches to the tympanum or the shoulder when hindlimbs adpressed forward towards body; (6) head length more than 30% of SVL. In addition, the new species can be separated from all other congeners by uncorrected genetic distances: 12S rRNA, 1.1%-2.1% of the 16S rRNA, and 7.5%- 11.5% of the COI.