[1] Eszterhazy Karoly Catholic Univ, Dept Zool, Eger, Hungary
[2] Inst Soil Sci Agrotecnol & Plant Protect Nikola P, Agr Acad, Sofia, Bulgaria
[3] Res Ctr Humanities, Inst Archaeogenom, Budapest, Hungary
[4] Univ Haifa, Inst Evolut, Haifa, Israel
[5] Eotvos Lorand Univ, Dept Systemat Zool & Ecol, Pazmany P Setany 1-C, H-1117 Budapest, Hungary