[1] Univ Calabria, Dept Biol Ecol & Earth Sci, Via P Bucci 4-B, I-87036 Arcavacata Di Rende, Italy
[2] Univ Naples Federico II, Dept Biol, Via Cinthia 26, I-80126 Naples, Italy
[3] Russian Acad Sci, Zool Inst, St Petersburg 190121, Russia
[4] CNR, Inst Biomed Technol, I-56124 Pisa, Italy