[1] Beijing Forestry Univ, Sch Soil & Water Conservat, Yanchi Res Stn, Beijing 100083, Peoples R China
[2] Beijing Forestry Univ, Key Lab State Forestry Grassland Adm Soil & Water, Beijing 100083, Peoples R China
[3] Beijing Forestry Univ, State Key Lab Efficient Prod Forest Resources, Beijing 100083, Peoples R China
[4] Utah State Univ, Dept Wildland Resources, Logan, UT 84321 USA
[5] Utah State Univ, Ecol Ctr, Logan, UT 84321 USA
[6] Chinese Acad Sci, Inst Appl Ecol, Erguna Forest Steppe Ecotone Res Stn, Shenyang 110016, Peoples R China
[7] Chinese Acad Sci, Inst Bot, State Key Lab Vegetat & Environm Change, Beijing 100093, Peoples R China
[8] Beijing Forestry Univ, Engn Res Ctr Forestry Ecol Engn, Minist Educ, Beijing 100083, Peoples R China