[1] Anhui Normal Univ, Coll Life Sci, Wuhu 241000, Peoples R China
[2] Chinese Acad Sci, Kunming Inst Bot, Key Lab Plant Divers & Biogeog East Asia, Kunming 65000, Peoples R China
[3] Senckenberg Res Inst, Dept Bot & Mol Evolut, D-60325 Frankfurt, Germany
[4] Univ Oslo, Nat Hist Museum, N-0318 Oslo, Norway