[1] Rhodes Univ, Ctr Biol Control, Dept Zool & Entomol, Makhanda Grahamstown, South Africa
[2] Univ Chester, Dept Biol Sci, Chester, England
[3] Univ Free State, Dept Zool & Entomol, Afromontane Res Unit, Phuthaditjhaba, South Africa
[4] Rhodes Univ, Ctr Biol Control, Dept Zool & Entomol, ZA-6139 Makhanda Grahamstown, Eastern Cape, South Africa