[1] Inst Ecol, Red Ecoetol, Xalapa, Mexico
[2] Univ Bosque, Fac Ingn, Grp Agua Salud & Ambiente, Bogota, Colombia
[3] Univ Nacl Autonoma Mexico, Dept Biol Evolut, Inst Ecol, Ciudad Univ, Mexico City, Mexico
[4] Univ Estadual Mato Grosso Do Sul, Lab Bioecol Scarabaeoidea Scaralab, Aquidauana, Brazil
[5] Univ Nacl Mis UNaM, Inst Biol Subtrop IBS, CONICET, Posadas, Argentina
[6] Div Ciencias Ambientales, CONACYT, IPICYT, San Luis Potosi, Mexico