[1] Grp Estudio Cactaceas & Vida Silvestre Occidente, AC, Comala 28454, Colima, Mexico
[2] Univ Nacl Autonoma Mexico, Jardin Bot, Inst Biol, Zona Deport S-N, Ciudad De Mexico 04510, Mexico
[3] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, Herbario Luz Maria Villarreal de Puga, Zapopan 45200, Jalisco, Mexico
[4] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, CONAHCYT Lab Nacl Identificac & Caracterizac Veget, Zapopan 45200, Jalisco, Mexico