[1] Univ Reg Cariri, Dept Quim Biol, Lab Zool, Campus Pimenta, CE, Brazil
[2] Univ Sao Paulo, Fac Med Vet & Zootecnia, Dept Med Vet Prevent & Saude Anim, Sao Paulo, Brazil
[3] Secretaria Estado Saude Sao Paulo, Dept Labs Especializados, Superintendencia Controle Endemias, Sao Paulo, Brazil
[4] Univ Wolverhampton, Fac Sci & Engn, Wulfruna St, Wolverhampton WV1 1LY, England
[5] Leibniz Inst Anal Biodiversitatswandels, Zent Taxon & Morphol, Sekt Herpetol, Museum Koenig, Adenauerallee 160, D-53113 Bonn, Germany
[6] Victor Valley Coll, Dept Biol, 18422 Bear Valley Rd, Victorville, CA 92395 USA