[1] Univ Lausanne, Ctr Integrat Genom, Lausanne, Switzerland
[2] Univ Lausanne, Dept Computat Biol, Lausanne, Switzerland
[3] Swiss Inst Bioinformat, Lausanne, Switzerland
[4] Univ Tartu, Inst Mol & Cell Biol, Tartu, Estonia
[5] Univ Tartu, Estonian Genome Ctr, Tartu, Estonia
[6] Univ Ctr Primary Care & Publ Hlth, Lausanne, Switzerland