[1] Normandie Univ, UMR BOREA 8067, MNHN, UPMC,IRD 207,UNICAEN,CNRS,UCBM, CS 14032, Caen, France
[2] France Energies Marines ITE EMR, 525 Ave Alexis de Rochon, F-29280 Plouzane, France
[3] Univ Paris, Lab Dynam Sociales & Recomposit Espaces, LADYSS, UMR 7533, F-75205 Paris 13, France
[4] Univ Brest, CNRS, IRD, Ifremer,LEMAR,IUEM, Rue Dumont dUrville,BP 70, F-29280 Plouzane, France
[5] Univ Littoral Cote dOpale, Univ Lille, CNRS, IRD,UMR LOG 8187,Lab Oceanol & Geosci, F-62930 Wimereux, France
[6] Normandie Univ, UNICAEN, CNRS, UMR M2C 6143, 24 Rue Tilleuls,CS 14032, Caen, France
[7] Normandie Univ, UNICAEN, CNRS, UMR LMNO 6139, 6 Bd Marechal Juin,CS 14032, Caen, France
[8] Ifremer, Unite Halieut Manche Mer Nord Ifremer, HMMN, F-62200 Boulogne Sur Mer, France
[9] Normandie Univ, UNICAEN, MRSH, CS 14032, Caen, France