[1] Sun Yat Sen Univ, Sch Marine Sci, Southern Marine Sci & Engn Guangdong Lab Zhuhai, Zhuhai 519000, Peoples R China
[2] Univ Macau, Dept Ocean Sci & Technol, State Key Lab Internet Things Smart City, Taipa 999078, Macau, Peoples R China
[3] Guangdong Prov Key Lab Marine Resources & Coastal, Guangzhou 510000, Peoples R China
[4] Minist Educ, Pearl River Estuary Marine Ecosyst Res Stn, Zhuhai 519000, Peoples R China