[1] GEOMAR Helmholtz Ctr Ocean Res Kiel, Marine Ecol Dept, Dusternbrooker Weg 20, D-24105 Kiel, Germany
[2] Leibniz Inst Sci & Math Educ IPN, Kiel Sci Factory, Bot Garten 16i, D-24118 Kiel, Germany
[3] Univ Kiel, Bot Garten 16i, D-24118 Kiel, Germany
[4] GEOMAR Helmholtz Ctr Ocean Res Kiel, Marine Geosyst, Wischhofstr 1-3, D-24148 Kiel, Germany
[5] Marine Ecol Dept, GEOMAR, Dusternbrooker Weg 20, D-24105 Kiel, Germany