[1] Univ Seville, Fac Biol, Dept Zool, Biodivers & Ecol Acaat, Seville, Spain
[2] Univ Seville, Dept Zool, Area Invest Biol I D I Acuario Sevilla, Lab Biol Marina,Fac Biol, Seville, Spain
[3] Univ Malaga, Grp Oceanog Fis, Inst Biotecnol & Desarrollo Azul IBYDA, Malaga, Spain
[4] Univ Seville, Fac Biol, Dept Zool, Av Reina Mercedes 6, Seville 41012, Spain