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Chinese Acad Agr Sci, Lanzhou Vet Res Inst, Key Lab Vet Publ Hlth, State Key Lab Vet Etiol Biol,Agr Minist, Lanzhou 730046, Gansu, Peoples R China Chinese Acad Agr Sci, Lanzhou Vet Res Inst, Key Lab Vet Publ Hlth, State Key Lab Vet Etiol Biol,Agr Minist, Lanzhou 730046, Gansu, Peoples R China