[1] Chinese Acad Sci, Inst Appl Ecol, Shenyang 110016, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[3] Chinese Acad Sci, Inst Appl Ecol, Key Lab Conservat Tillage & Ecol Agr, Shenyang 110016, Liaoning, Peoples R China
[4] Dalian Minzu Univ, Coll Environm & Resources, Dalian 116600, Peoples R China