[1] Obihiro Univ Agr & Vet Med, Grad Sch Anim & Vet Sci & Agr, 2 11 Inada, Obihiro, Hokkaido 0808555, Japan
[2] Jomo Kenyatta Univ Agr & Technol, Dept Med Lab Sci, Coll Hlth Sci, POB 62000 00200, Nairobi, Kenya
[3] Obihiro Univ Agr & Vet Med, Dept Vet Med, 2 11 Inada, Obihiro, Hokkaido 0808555, Japan
[4] Obihiro Univ Agr & Vet Med, Res Ctr Global Agromed, 2 11 Inada, Obihiro, Hokkaido 0808555, Japan
[5] Obihiro Univ Agr & Vet Med, Natl Res Ctr Protozoan Dis, 2 11 Inada, Obihiro, Hokkaido 0808555, Japan
[6] Tokyo Univ Agr & Technol, Ctr Infect Dis Epidemiol & Prevent Res CEPiR, Tokyo, 1838509, Japan