[1] Chinese Acad Sci, Inst Zool, Beijing 100101, Peoples R China
[2] Langfang Normal Univ, Coll Life Sci, Langfang 065000, Hebei, Peoples R China
[3] Tokyo Metropolitan Univ, Grad Sch Sci, Dept Biol Sci, Systemat Zool Lab, 1-1 Minami Osawa, Hachioji, Tokyo 1920397, Japan
[4] Museo Storia Nat Verona, Dept Zool, Lungadige Porta Vittoria 9, I-37129 Verona, Italy
[5] Vietnam Acad Sci & Technol VAST, Vietnam Natl Museum Nat VNMN, 18 Hoang Quoc Viet,Cau Giay, Hanoi, Vietnam