New Discovery of Hipparion theobaldi Skull from the Late Miocene of Padhri, District Jhelum, Punjab, Pakistan and Associated Mammalian Fossil Assemblage

A fossil-bearing locality near Padhri village, Dhok Pathan, 55 km away from the tehsil Dina, Jhelum District, in the Potwar Plateau, Middle Siwaliks, Punjab, northern Pakistan, is significantly rich in mammalian fossils. This site has provided an abundant mammalian fossil fauna of Late Miocene age from the Dhok Pathan Formation(Fm.). The recovered material belongs to four families: Equidae(horses), Rhinocerotidae(rhinos), Bovidae(cows), and Suidae(pigs). We discovered a new skull of hipparionine Hipparion theobaldi from this locality along with 22 specimens from the associated assemblage of fossil mammals. The recovered material includes seven other species: the aceratheriine Chilotherium intermedium, boselaphines Tragoportax punjabicus, Selenoportax vexillarius, Pachyportax latidens, the antelope Gazella lydekkeri and suinine Propotamochoerus hysudricus. The specimens are isolated teeth, fragments of maxilla, mandibles and horn cores. The Dhok Pathan Fm. is generally composed of claystone, siltstone and sandstone beds and, based on the mammalian fauna, the Padhri fossil locality is dated as Late Miocene. Thi99 s formation was deposited in a subtropical paleoenvironment and the predominance of fossil bovids indicates extremely moist conditions with small but frequent standing water bodies.