[1] Nat Hist Museum, Life Sci Dept, London SW7 5BD, England
[2] Univ Gothenburg, Dept Marine Sci, Box 463, S-40530 Gothenburg, Sweden
[3] Gothenburg Global Biodivers Ctr, Box 463, S-40530 Gothenburg, Sweden
[4] Univ Southampton, Sch Ocean & Earth Sci, Southampton SO14 3ZH, England
[5] NORCE Norwegian Res Ctr, POB 22 Nygardsgaten, NO-5838 Bergen, Norway