[1] Tribhuvan Univ, Cent Dept Zool, Kirtipur 44613, Nepal
[2] Sukuna Multiple Campus, Sundarharaincha 56604, Nepal
[3] Small Mammals Conservat & Res Fdn, Balkhu 44614, Nepal
[4] Hungarian Nat Hist Museum, Dept Zool, H-1088 Budapest, Hungary
[5] Hungarian Univ Agr & Life Sci, Inst Agron, H-2100 Godollo, Hungary
[6] Eszterhazy Karoly Catholic Univ, Dept Zool, H-3300 Eger, Hungary
[7] Guangzhou Univ, Sch Life Sci, Guangzhou 510006, Peoples R China
[8] Univ Pecs, Szentagothai Res Ctr, Natl Lab Virol, H-7624 Pecs, Hungary