[1] Univ Coimbra, Ctr Funct Ecol, Dept Life Sci, Coimbra, Portugal
[2] Univ Mons, Res Inst Biosci, Lab Zool, Mons, Belgium
[3] Tagis Ctr Conservacao Borboletas Portugal, Avis, Portugal
[4] Univ Coimbra, Sci Museum, Coimbra, Portugal
[5] Nat Biodivers Ctr, Leiden, Netherlands