[1] South African Natl Biodivers Inst, Natl Herbarium, Private Bag X101, ZA-0001 Pretoria, South Africa
[2] Univ Western Cape, Dept Biodivers & Conservat Biol, Private Bag X17, ZA-7535 Cape Town, South Africa
[3] South African Natl Biodivers Inst, Kirstenbosch Res Ctr, Compton Herbarium, Rhodes Dr, ZA-7700 Cape Town, South Africa
[4] Univ Johannesburg, Dept Bot & Plant Biotechnol, POB 524,Auckland Park, ZA-2006 Johannesburg, South Africa
[5] Univ KwaZulu Natal, Sch Biol & Conservat Sci, Res Ctr Plant Growth & Dev, Private Bag X01, ZA-3209 Pietermaritzburg, South Africa