[1] IST Austria, A-3400 Klosterneuburg, Austria
[2] Max Planck Inst Evolutionary Biol, Max Planck Res Grp Dynam Social Behav, D-24306 Plon, Germany
[3] Queensland Univ Technol, Sch Math Sci, Brisbane, Qld 4000, Australia
[4] Univ Queensland, Sch Econ, Brisbane, Qld 4067, Australia