[1] Univ Parakou, Fac Agron, Lab Ecol Bot & Biol Vegetale, 03 BP 123, Parakou, Benin
[2] Univ Liege, Fac Sci, Dept Sci & Gest Environm, Lab Eau Environm Dev,t,Arlon CampusEnvironm,UR SPH, 185 Ave Longwy, B-6700 Arlon, Belgium
[3] Univ Parakou, Fac Agron, Lab Etud & Rech Forestieres LERF, 03 BP 123, Parakou, Benin